Taught at the Universidad Costa Rica every two years in spring semesters
Limnology is the study of inland waters - lakes, streams, wetlands, and reservoirs. I teach this class jointly with Professor Gerardo Umaña at UCR. Topics that we cover include: the origin of lakes, the importance of chemical and physical properties, biogeochemical cycles (e.g., carbon, nitrogen, phosphorus, iron and silicon), freshwater pollution and eutrophication, food-webs and energy flow, and paleolimnology. The course also includes training in chemical analysis and water sampling techniques, as well as species identification of phytoplankton, zooplankton, macroinvertebrates, and fish. During a weekend field trip, students get practical experience collecting samples in lake ecosystems.